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  • A panning exposure captures the motion of a Masai ostrich (Struthio camelus massaicus) walking through tall grass in the savannah of the Maasai Mara National Reserve in Kenya. The ostrich is the largest living bird species and can reach a height of 9 feet, 2 inches (2.8 meters) — about half of the height coming from its neck. The ostrich cannot fly, but can run faster than any other bird, reaching a top speed of 40 miles per hour (70 kilometers per hour).
    Kenya_Maasai-Mara_Ostrich_Panning_61...jpg
  • A greater yellowlegs (Tringa melanoleuca) walks in the shallow, muddy water of the Stillaguamish River near Stanwood, Washington. Greater yellowlegs feed on insects, small fish, marine worms, and crustaceans, sometimes using their bills to stir up water.
    Yellowlegs-Greater_Walking_Stillagua...jpg
  • An African lion (Panthera leo) walks through tall savannah grass in the golden light of sunrise in the Maasai Mara National Reserve, Kenya.
    Kenya_Maasai-Mara_Lion_Grass_3503.jpg
  • An American bison (Bison bison) walks in deep snow near Fountain Flat in Yellowstone National Park, Wyoming. Bison are well equiped for harsh winter conditions. They grow a winter coat of wooly underfur, which has coarse hairs that protect them from the elements. The humps on their backs also contain muscles supported by long vertebrae that help swing their heads to move vast amounts of snow.
    Bison_Snow_Yellowstone_4420.jpg
  • A Masai giraffe (Giraffa camelopardalis tippelskirchi) walks in the savanna of the Maasi Mara National Reserve in Kenya as yellow-billed oxpeckers (Buphagus africanus) gather on its back.
    Kenya_Maasai-Mara_Giraffe_Walking_40...jpg
  • A long exposure captures the motion of a cheetah (Acinonyx jubatus) walking through tall grass in the Maasai Mara National Reserve, Kenya.
    Kenya_Maasai-Mara_Cheetah_Walking_Pa...jpg
  • A couple, rendered in silhouette, walk past a wind-swept tree on a bluff overlooking the Pacific Ocean at Ecola State Park near Cannon Beach, Oregon.
    WindSweptTreeCouple.jpg
  • A long exposure captures the motion of an African lion (Panthera leo) walking through tall savannah grass in the golden light of sunrise in the Maasai Mara National Reserve, Kenya.
    Kenya_Maasai-Mara_Lion_Walking_Panni...jpg
  • A long exposure captures the motion of a cheetah (Acinonyx jubatus) walking through tall grass in the Maasai Mara National Reserve, Kenya.
    Kenya_Maasai-Mara_Cheetah_Walking_Pa...jpg
  • A long exposure captures the motion of a leopard (Panthera pardus) walking through tall grass in the Maasai Mara National Reserve, Kenya.
    Kenya_Maasai-Mara_Leopard_Walking_Pa...jpg
  • A large moose (Alces alces) walks in deep snow in search for food in Yellowstone National Park, Wyoming. Moose, which are known as elk in Eurasia, have thick skin and other features that make them well-adapted for cold weather.
    Moose_Snow_Walking_Yellowstone_0888.jpg
  • A coyote (Canis latrans) walks among the golden grasses in a snow-covered field in search of food in Yellowstone National Park, Montana.
    Coyote_Snowy-Field_Yellowstone_8084.jpg
  • Four yellow-eyed penguins (Megadyptes antipodes), also known as Hoiho, walk across the beach at Jack's Bay, located in the Catlins on the southern tip of the South Island of New Zealand. Yellow-eyed penguins are endangered and are one of the most rare penguins in the world with a total population of only about 4,000. About 90 percent of the yellow-eyed penguin's diet consists of fish. During the breeding season, many of the penguins spend the entire day hunting in the ocean. They enter the Pacific Ocean at dawn and return at dusk, venturing as far as 25 kilometers (16 miles) offshore and diving to depts of up to 120 meters (394 feet).
    NZ_Penguins_YellowEyed_JacksBay_8291.jpg
  • A Mojave Desert Tortoise (Gopherus agassizii) walks near shrubs in the Desert Tortoise Natural Area in California City, California.
    Desert-Tortoise-Walking_7815.jpg
  • An African lion (Panthera leo) walks through tall savannah grass in the golden light of sunrise in the Maasai Mara National Reserve, Kenya.
    Kenya_Maasai-Mara_Lion_Grass_3498.jpg
  • A long exposure captures the motion of an African lion (Panthera leo) walking through tall savannah grass in the golden light of sunrise in the Maasai Mara National Reserve, Kenya.
    Kenya_Maasai-Mara_Lion_Walking_Panni...jpg
  • A Masai ostrich (Struthio camelus massaicus) walks through tall grass in the savannah of the Maasai Mara National Reserve in Kenya. The ostrich is the largest living bird species and can reach a height of 9 feet, 2 inches (2.8 meters) — about half of the height coming from its neck. The ostrich cannot fly, but can run faster than any other bird, reaching a top speed of 40 miles per hour (70 kilometers per hour).
    Kenya_Maasai-Mara_Ostrich_Walking_61...jpg
  • An American bison (Bison bison) forces its way through deep snow near Fountain Flat in Yellowstone National Park, Wyoming. Bison are well equipped for harsh winter conditions. They grow a winter coat of woolly underfur, which has coarse hairs that protect them from the elements. The humps on their backs also contain muscles supported by long vertebrae that help swing their heads to move vast amounts of snow.
    Bison_Snow_Yellowstone_4275.jpg
  • An American bison (Bison bison) forces its way through deep snow near Fountain Flat in Yellowstone National Park, Wyoming. Bison are well equiped for harsh winter conditions. They grow a winter coat of wooly underfur, which has coarse hairs that protect them from the elements. The humps on their backs also contain muscles supported by long vertebrae that help swing their heads to move vast amounts of snow.
    Bison_Snow_Yellowstone_4275sq.jpg
  • A young family walks on the beach near Haystack Rock, Oregon, in thick fog. Haystack Rock, remnants of an ancient lava flow, is one of the world's largest monoliths.
    HaystackFogFamily.jpg
  • Backed by the colors of sunrise, an African lion (Panthera leo) walks atop a small ridge in the the savannah of the Maasai Mara National Refuge in Kenya.
    Kenya_Maasai-Mara_Lion_Sunrise_Panor...jpg
  • A pronghorn (Antilocapra americana) walks on a snow-dusted hill in Yellowstone National Park, Montana. They are often mistakenly referred to as antelope, although they are more closely related to giraffes than to Old World antelope.
    Pronghorn_Winter_Yellowstone_3328.jpg
  • A bull elk (Cervus canadensis) walks in a snow-covered field near Madison in Yellowstone National Park, Wyoming.
    Elk_Yellowstone_Snow-Storm_4050.jpg
  • A captive cougar walks through a dense forest in Washington state.
    CougarInForest.jpg
  • In the golden evening light, a plains zebra (Equus quagga) walks in tall grass of the savanna in the Maasai Mara National Reserve in Kenya.
    Kenya_Maasai-Mara_Zebra_Acacia-Tree_...jpg
  • A long exposure captures the motion of topis (Damaliscus lunatus jimela) walking through tall grass in the savannah of the Maasai Mara National Reserve in Kenya.
    Kenya_Maasai-Mara_Topis_Panning_5112.jpg
  • A greater yellowlegs (Tringa melanoleuca) casts its shadow and a reflection as it walks along the muddy banks of the Stillaguamish River at sunset on Leque Island near Stanwood, Washington.
    Yellowlegs-Greater_Shadow_Eide-Road_...jpg
  • A bighorn sheep (Ovis canadensis) walks on a snowy ridge that borders the Lamar Valley in Yellowstone National Park, Wyoming.
    Bighorn-Sheep_Ridge_Winter_Yellowsto...jpg
  • A cheetah (Acinonyx jubatus) tries to blend in with the golden savannah grass as it looks for prey in the Maasai Mara National Reserve in Kenya.
    Kenya_Maasai-Mara_Cheetah_Golden-Gra...jpg
  • A black bear (Ursus americanus) leads her cub through a field of blueberries in Waterton Lakes National Park, Alberta, Canada. It is the most common bear species in North America, found in 41 of the 50 U.S. states and all Canadian provinces except Prince Edward Island.
    BlackBearWalking.jpg
  • A group of yellow-eyed penguins (Megadyptes antipodes), also known as Hoiho, preen themselves as they march up the hillside to their colony at Nugget Point in the Catlins at the southern tip of the South Island of New Zealand. Yellow-eyed penguins are endangered and are one of the most rare penguins in the world with a total population of only about 4,000. About 90 percent of the yellow-eyed penguin's diet consists of fish. During the breeding season, many of the penguins spend the entire day hunting in the ocean. They enter the Pacific Ocean at dawn and return at dusk, venturing as far as 25 kilometers (16 miles) offshore and diving to depts of up to 120 meters (394 feet).
    NZ_Penguins_YellowEyed_NuggetPoint_7...jpg
  • A moose (Alces alces) trudges through deep snow near Pebble Creek in Yellowstone National Park, Wyoming. Moose are known as elk in Eurasia and are the largest and heaviest living members of the deer family. Males, known as bulls, grow fresh antlers each year, dropping them in the winter.
    Moose_Winter_Yellowstone_8786.jpg
  • A moose (Alces alces) trudges through deep snow near Pebble Creek in Yellowstone National Park, Wyoming. Moose are known as elk in Eurasia and are the largest and heaviest living members of the deer family. Males, known as bulls, grow fresh antlers each year, dropping them in the winter.
    Moose_Winter_Yellowstone_8751.jpg
  • A greater yellowlegs (Tringa melanoleuca) casts its reflection on shallow water on a foggy morning on Leque Island near Stanwood, Washington.
    Yellowlegs-Greater_Foggy-Reflection_...jpg
  • A yellow-eyed penguin (Megadyptes antipodes), also known as Hoiho, waits for members of its colony to return at the edge of the Pacific Ocean at Nugget Point in the Catlins at the southern tip of the South Island of New Zealand. Yellow-eyed penguins are endangered and are one of the most rare penguins in the world with a total population of only about 4,000. About 90 percent of the yellow-eyed penguin's diet consists of fish. During the breeding season, many of the penguins spend the entire day hunting in the ocean. They enter the Pacific Ocean at dawn and return at dusk, venturing as far as 25 kilometers (16 miles) offshore and diving to depts of up to 120 meters (394 feet).
    NZ_Penguin_YellowEyed_NuggetPoint_74...jpg
  • Three yellow-eyed penguins (Megadyptes antipodes), also known as Hoiho, march up the hillside to their colony at Nugget Point in the Catlins at the southern tip of the South Island of New Zealand. Yellow-eyed penguins are endangered and are one of the most rare penguins in the world with a total population of only about 4,000. About 90 percent of the yellow-eyed penguin's diet consists of fish. During the breeding season, many of the penguins spend the entire day hunting in the ocean. They enter the Pacific Ocean at dawn and return at dusk, venturing as far as 25 kilometers (16 miles) offshore and diving to depts of up to 120 meters (394 feet).
    NZ_Penguins_YellowEyed_NuggetPoint_7...jpg
  • Two bison (Bison bison) trudge through deep snow near Fountain Flat in Yellowstone National Park, Wyoming. Bison are well equipped for harsh winter conditions. They grow a winter coat of woolly underfur, which has coarse hairs that protect them from the elements. The humps on their backs also contain muscles supported by long vertebrae that help swing their heads to move vast amounts of snow.
    Bison_Snow_Two_Fountain-Flat_Yellows...jpg
  • A moose (Alces alces) trudges through deep snow near Pebble Creek in Yellowstone National Park, Wyoming. Moose are known as elk in Eurasia and are the largest and heaviest living members of the deer family. Males, known as bulls, grow fresh antlers each year, dropping them in the winter.
    Moose_Winter_Yellowstone_8914.jpg
  • Chincoteague ponies (Equus caballus), also known as Assateague horses, walk through a marsh on Assateague Island in the Chincoteague National Wildlife Refuge in Virginia. Chincoteague ponies are small — typically 12-13 hands (about 4 feet tall) — their growth stunted by the limited food and harsh environment of Assateague Island. About 300 wild — technically feral — ponies roam the island on the Atlantic coast. There is some dispute as to how the ponies ended up on the island. Some researchers believe the ponies are survivors of the wreck of a Spanish galleon, La Galga, which sank just off the coast in 1750; the U.S. Fish and Wildlife Service believes they are descendants of horses owned by early colonial settlers.
    Assateague-Island_Chincoteague-Ponie...jpg
  • Two bighorn sheep (Ovis canadensis) walk down the flank of Druid Peak in Yellowstone National Park, Wyoming.
    Sheep-Bighorn_Yellowstone_5748.jpg
  • A pair of American bison (Bison bison) walk down the edge of a forested hillside into a meadow. Bison are also known as American buffalo.
    Bison_Pair-On-Forested-Hillside_4086.jpg
  • Thousands of snow geese walk and fly in formation over a farmer's field near Mount Vernon, Washington. More than 30,000 snow geese winter in the area, feeding on grass and other plants before flying north for the summer.
    SnowGeeseLinedUp.jpg
  • Two sandhill cranes (Grus canadensis) are rendered in silhouette as they walk across a pond in Bosque del Apache, New Mexico, at sunrise.
    SandhillCranes_Pond_Silhouette_7524.jpg
  • A coyote (Canis latrans) walks on the frozen Yellowstone River to hunt for prey in Yellowstone National Park, Wyoming.
    Coyote_Yellowstone-River_Frozen_9288.jpg
  • A coyote (Canis latrans) walks on the frozen Yellowstone River to hunt for prey in Yellowstone National Park, Wyoming.
    Coyote_Yellowstone-River_Frozen_9316.jpg
  • A raccoon (Procyon lotor) carefully walks out onto a floating log to forage in Lake Washington in Renton, Washington.
    Raccon_Log_Lake-Washington_Renton_49...jpg
  • A coyote (Canis latrans) walks on an ice-covered road as bison graze in the distance in the Lamar Valley of Yellowstone National Park, Wyoming.
    Coyote_Bison_Snow_Yellowstone_8036.jpg
  • A Chincoteague pony (Equus caballus), also known as an Assateague horse, walks through a marsh on Assateague Island in the Chincoteague National Wildlife Refuge in Virginia. About 300 wild — technically feral — ponies roam the island on the Atlantic coast. There is some dispute as to how the ponies ended up on the island. Some researchers believe the ponies are survivors of the wreck of a Spanish galleon, La Galga, which sank just off the coast in 1750; the U.S. Fish and Wildlife Service believes they are descendants of horses owned by early colonial settlers.
    Assateague-Island_Chincoteague-Pony_...jpg
  • An African savanna elephant (Loxodonta africana) walks in the savannah of the Maasai Mara National Reserve in Kenya as seen from a hot air balloon in early morning light.
    Kenya_Maasai-Mara_Elephant_Aerial_33...jpg
  • A raccoon (Procyon lotor) carefully walks out onto a floating log to forage in Lake Washington in Renton, Washington.
    Raccon_Log_Lake-Washington_Renton_48...jpg
  • A coyote (Canis latrans) walks on the frozen Yellowstone River to hunt for prey in Yellowstone National Park, Wyoming.
    Coyote_Yellowstone-River_Frozen_9271.jpg
  • A raccoon (Procyon lotor) carefully walks out onto a floating log to forage in Lake Washington in Renton, Washington.
    Raccon_Log_Lake-Washington_Renton_48...jpg
  • A young red fox (Vulpes vulpes) kit walks in the colorful prairie of the San Juan Island National Historical Park on San Juan Island, Washington. All of the foxes in the park are technically red foxes, even if they are black, silver or tan. Red foxes were introduced to San Juan Island on various occasions in the 1900s.
    Fox-Red_Kit_Black_Prairie_San-Juan_4...jpg
  • On a cold morning in the Maasai Mara National Reserve in Kenya, an African lion's (Panthera leo) breath is visible as golden mist as it walks through tall grass on the savannah.
    Kenya_Maasai-Mara_Lion_Grass_Breath_...jpg
  • A red fox (Vulpes vulpes) walks through snow to hunt in the Custer Gallatin National Forest near Cooke City, Montana.
    Fox-Red_Hunting_Winter_Gallatin-NF_7...jpg
  • A North American river otter (Lontra canadensis) walks on ice covering the Yellowstone River next to a patch of open water in Yellowstone National Park, Wyoming.
    Otter-River_Yellowstone-River_Frozen...jpg
  • A wild turkey (Meleagris gallopavo) walks through a grassy field in the Blue Mountains of Washington state.
    Turkey_Blue-Mountains-WA_8707.jpg
  • An Atlantic puffin (Fratercula arctica), right, walks up to check out a razorbill (Alca torda) on the Látrabjarg bird cliff in western Iceland. Látrabjarg is Europe's largest bird cliff: 14 km (8.7 miles) long and standing up to 440 meters (1444 feet) above the Atlantic Ocean. During the breeding season, Látrabjarg hosts as much as 40 percent of the breeding populations of some birds, including razorbills.
    Razorbill_Puffin_Latrabjarg_2375.jpg
  • A black-tailed deer (Odocoileus hemionus) walks through a field of summer wildflowers in the Paradise Meadows of Mount Rainier National Park, Washington.
    RainierParadise_Deer_3735.jpg
  • An African lion (Panthera leo) cub walks in the savannah grass as several lionesses rest in the Maasai Mara National Reserve in Kenya.
    Kenya_Maasai-Mara_Lion_Pride_Cub_865...jpg
  • Bathed in the golden light of sunrise, an African lion (Panthera leo) walks through tall grass in the savannah of the Maasai Mara National Refuge in Kenya.
    Kenya_Maasai-Mara_Lion_Golden-Grass_...jpg
  • A mustang (Equus caballus) walks under a canopy of mesquite trees in the Tonto National Forest in the woods near the Salt River in Arizona. The free-roaming mustangs are commonly referred to as wild horses, but they are descendants of domesticated Spanish horses that were brought to the continent in the 16th century.
    Mustang_Forest_Coon-Bluff_4040.jpg
  • An American pipit (Anthus rubescens) poses with an insect that it caught above the tree line in Mount Rainier National Park, Washington. American pipits are found in the tundra and on alpine slopes and they forage by walking on the ground, taking insects from the ground or from low plants.
    Pipit-American_Insect_RainierNP_4437.jpg
  • A killdeer (Charadrius vociferus) walks around the edge of Kendall Lake in the Cuyahoga Valley National Park, Ohio.
    OH_Cuyahoga-Valley-NP_Kendall-Lake_K...jpg
  • An arctic fox (Vulpes lagopus) in transition from its winter to summer coat walks across on old lava field on Iceland's Snæfellsnes peninsula. Arctic foxes have limbs, snouts and ears that are shorter than average, a feature that minimizes heat loss and the risk of frostbite. The pads of their feet are also covered in fur. Arctic foxes are primarily nocturnal; this fox was photographed shortly after midnight.
    Iceland_ArcticFox_7346.jpg
  • A red fox (Vulpes vulpes) walks along a dried creek bed near Swiftcurrent Creek in Glacier National Park, Montana.
    RedFox_Glacier.jpg
  • Three white-faced ibis (Plegadis chihi) feed at the edge of a pond in the Malheur National Wildlife Refuge in southeastern Oregon. Ibises forage in groups by walking slowly with their heads down, probing the mud with their long, curved bills.
    Ibis_WhiteFaced_Malheur_5334.jpg
  • A coyote (Canis latrans) walks out onto the frozen Crowley Lake near Mammoth Lakes, California.
    Coyote_CrowleyLake_Frozen_6799.jpg
  • An African lion (Panthera leo) licks its chops as it walks through tall grass in the savannah of the Maasai Mara National Refuge in Kenya.
    Kenya_Maasai-Mara_Lion_Lick_3478.jpg
  • A young red fox (Vulpes vulpes) kit walks in the colorful prairie of the San Juan Island National Historical Park on San Juan Island, Washington. Red foxes were introduced to San Juan Island on various occasions in the 1900s.
    Fox_Red_Kit_Prairie_San-Juan_4588.jpg
  • A red rock crab (Cancer productus) walks across the murky bottom of Puget Sound off Des Moines Beach Park, Des Moines, Washington.
    Crab_RedRock_DesMoines_F0112.jpg
  • A young red fox (Vulpes vulpes) kit walks in the colorful prairie of the San Juan Island National Historical Park on San Juan Island, Washington. All of the foxes in the park are technically red foxes, even if they are black, silver or tan. Red foxes were introduced to San Juan Island on various occasions in the 1900s.
    Fox-Red_Kit_Black_Prairie_San-Juan_6...jpg
  • A Plains bison (Bison bison) walks in a heavy snow storm in a field near the Madison River in Yellowstone National Park, Wyoming. American bison are sometimes mistakenly referred to as buffalo; they are only distantly related to true buffalo.
    Bison_Yellowstone_Snow-Storm_3907.jpg
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